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Extra info for Evolution of the Metazoan life cycle: a comprehensive theory
It seems possible to me that the formation of the mantle at the time when it was shifted to the larva was effected just below the already existing prototroch, which thus became pushed to its margin-to some degree a parallel to the conditions at the formation of the velum in the larvae of molluscs. 65 None of these differences is sufficiently fundamental that a relatively close relationship between the three groups of tentaculates could be questioned. By explaining certain of the previously very puzzling differ ences the present examination has on the contrary provided further support for such a relationship.
G. in the pelagic genus Pedalia (Fig. 14) and in the benthic family Flosculariidae (Melicertidae) . How is this agreement with an original trochophora to be understood ? There are only two possibilities. Either we are faced by a phylogenetically conditioned similarity, thus a similarity which can be traced back to con ditions in the common ancestral form of both rotifers and trochophora groups, or else this is nothing more than a phenomenon of convergence. According to the prevailing opinion the latter is the case.
It seems obvious that for the purposes of comparison we can disregard these structures. As far as the tentacles are concerned, we have already found it probable that the occurrence of this adult character in the actinotrocha is due to relatively late adultation. The transference of the tentacles to the larva is a process that in this connection is totally irrelevant. It need not necessarily have taken place. ) The ring of cilia, 59 SPECIAL SECTION TENTACULATA on the other hand, is a purely larval (adaptive pelagic) character and as such · cannot reasonably interfere with the comparison made.